1. Cell Biology, Neurobiology and Anatomy, Medical College of Wisconsin, Milwaukee, Wisconsin, United States
2. Department of Ophthalmology and Visual Sciences, Medical College of Wisconsin, Milwaukee, Wisconsin, United States
3. Morgridge Institute for Research, Madison, Wisconsin, United States
4. Department of Biology, University of Wisconsin Oshkosh, Oshkosh, Wisconsin, United States
5. Departments of Ophthalmology & Visual Sciences and Anatomical Sciences & Neurobiology, University of Louisville, Louisville, Kentucky, United States
6. Nuffield Laboratory of Ophthalmology, University of Oxford, Oxford, United Kingdom