1. Department of Ophthalmology, Kings College London, London, United Kingdom
2. KCL Department of Twin Research and Genetic Epidemiology, London, United Kingdom
3. Institute of Ophthalmology, University College London, London, United Kingdom
4. NIHR Biomedical Research Centre, Moorfield's Eye Hospital NHS Foundation Trust and UCL Institute of Ophthalmology, London, United Kingdom
5. Department of Public Health and Primary Care, Institute of Public Health, University of Cambridge School of Clinical Medicine, Cambridge, United Kingdom