1. Department of Ophthalmology and Visual Sciences, Carver College of Medicine, The University of Iowa, Iowa City, IA, United States
2. Center for the Prevention and Treatment of Visual Loss, Iowa City VA Healthcare System, Department of Veterans Affairs, Iowa City, IA, United States
3. The Department of Molecular Physiology and Biophysics, Carver College of Medicine, The University of Iowa, Iowa City, IA, United States