1. Department of Ophthalmology, Oita University, Oita, Japan
2. Laboratory for Retinal Regeneration, RIKEN Center for Developmental Biology, Kobe, Japan
3. Center for Stem Cell and Regenerative Medicine, Tokyo Medical and Dental University, Tokyo, Japan
4. Department of Ophthalmology & Visual Science, Tokyo Medical and Dental University, Graduate School of Medicine and Dental Sciences, Tokyo, Japan
5. Department of Ophthalmology & Visual Science, Tokyo Medical and Dental University, Graduate School of Medicine and Dental Sciences, Tokyo, Japan 5Miyata Eye Hospital, Miyakonojo, Japan