1. Department of Ophthalmology & Visual Sciences, Washington University School of Medicine, St. Louis, Missouri, United States 2Neuroscience Graduate Program, Division of Biology and Biomedical Sciences, Washington University School of Medicine, St. Louis, Missouri, United States
2. College of Optometry, University of Missouri–St. Louis, St. Louis, Missouri, United States
3. Department of Ophthalmology & Visual Sciences, Washington University School of Medicine, St. Louis, Missouri, United States 4Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States 5Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States