1. Graduate Department of Neuroscience, Cumming School of Medicine, University of Calgary, Calgary, AB, Canada
2. Department of Biochemistry and Molecular Biology, University of Calgary, Calgary, AB, Canada
3. Department of Biology, University of Victoria, Victoria, BC, Canada
4. Department of Medical Genetics, Cumming School of Medicine, University of Calgary, Calgary, AB, Canada
5. Department of Molecular Physiology and Biophysics, Department of Otolaryngology Head-Neck Surgery and Department of Neurology, University of Iowa, Iowa City, IA, USA
6. Department of Ophthalmology and Visual Sciences, Faculty of Medicine, and Clinical Vision Sciences Program, Faculty of Health Dalhousie University, NS, Canada
7. Department of Cell Biology and Anatomy and Department of Surgery, Hotchkiss Brain Institute, and Alberta Children's Hospital Research Institute, Cumming School of Medicine, University of Calgary, Calgary, AB, Canada
8. Department of Medical Genetics, and Department of Surgery, Alberta Children's Hospital Research Institute, and Hotchkiss Brain Institute, Cumming School of Medicine, University of Calgary, Calgary, AB, Canada