1. From the School of Mathematical Sciences, University of Nottingham, Nottingham, United Kingdom; the2Centre for Plant Integrative Biology, University of Nottingham, Sutton Bonington, United Kingdom; the
2. School of Mathematical Sciences and the4Institute of Health and Biomedical Innovation, Queensland University of Technology, Brisbane, Queensland, Australia; the
3. School of Mathematical Sciences, University of Southampton, Southampton, United Kingdom; the
4. From the School of Mathematical Sciences, University of Nottingham, Nottingham, United Kingdom; the
5. Department of Bioengineering, Imperial College London, London, United Kingdom; and the
6. Department of Ophthalmology, Queen’s Medical Centre, Nottingham, United Kingdom.