1. From the The Jackson Laboratory, Bar Harbor, Maine; the
2. Departments of Ophthalmology and Visual Sciences and
3. Departments of Ophthalmology and Visual Sciences and3Cell and Developmental Biology, University of Michigan, Ann, Arbor, Michigan; and the
4. Scheie Eye Institute and Kirby Center for Molecular Ophthalmology, University of Pennsylvania, Philadelphia, Pennsylvania; the
5. Jules Stein Eye Institute, University of California at Los Angeles, Los Angeles, California.