1. Department of Ophthalmology & Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States
2. Department of Pathology and Molecular Medicine, McMaster University, Hamilton, Ontario, Canada
3. Department of Pediatrics, Wayne State University, Detroit, Michigan, United States
4. Department of Craniofacial Biology and Department of Cell and Developmental Biology, University of Colorado–Denver, Denver, Colorado, United States
5. Department of Ophthalmology & Visual Sciences, University of Michigan, Ann Arbor, Michigan, United States 5Department of Cell & Developmental Biology, University of Michigan, Ann Arbor, Michigan, United States