1. Department of Anatomy and Cell Biology Wayne State University School of Medicine, Detroit, Michigan, United States 2Department of Ophthalmology, Wayne State University School of Medicine, Detroit, Michigan, United States
2. Department of Medicine, School of Medicine, Case Western Reserve University, Cleveland, Ohio, United States
3. Department of Anatomy and Cell Biology Wayne State University School of Medicine, Detroit, Michigan, United States
4. Department of Ophthalmology and Visual Sciences, Washington University School of Medicine, St. Louis, Missouri, United States