1. From the Centre for Vision Research, Department of Ophthalmology and the Westmead Millennium Institute, the
2. School of Applied Vision Sciences, Faculty of Health Sciences, University of Sydney, Sydney; the
3. ARC Centre of Excellence in Vision Science and Research School of Biological Sciences, Australian National University, Canberra, Australia; the
4. Department of Preventive Medicine, and the5Doheny Eye Institute and Department of Ophthalmology, Keck School of Medicine, University of Southern California, Los Angeles, California.