1. Department of Cell Biology, National Institute for Basic Biology, Okazaki 444-8585, Japan
2. Department of Molecular Biomechanics, School of Life Science, The Graduate University for Advanced Studies, Okazaki 444-8585, Japan
3. Department of Biosciences, High Technology Research Center, Teikyo University of Science and Technology, Yamanashi 409-0193, Japan
4. PRESTO, Japan Science and Technology Corporation, Kawaguchi 332-0012, Japan
5. Division of Organic Chemistry, Institute for Protein Research, Osaka University, Suita 565-0871, Japan