1. Tsukita Cell Axis Project, Exploratory Research for Advanced Technology, Japan Science and Technology Corporation, Kyoto Research Park, Shimogyo-ku, Kyoto 600-8813, Japan
2. Department of Cell Biology, Kyoto University Faculty of Medicine, Sakyo-ku, Kyoto 606-8501, Japan
3. Department of Dermatology, Osaka University School of Medicine, Suita 565-0871, Japan
4. Laboratory of Cell Biology, KAN Research Institute Inc., Kyoto Research Park, Shimogyo-ku, Kyoto 600-8815, Japan
5. Department of Molecular Cell Biology, Institute of DNA Medicine, Jikei University School of Medicine, Minato-ku, Tokyo 105-0003, Japan