1. School of Biological Sciences, Faculty of Biology, Medicine and Health, University of Manchester 1 , Manchester, UK
2. A*STAR Institute of Medical Biology 2 , Singapore, Singapore
3. MRC Laboratory of Molecular Biology, Francis Crick Avenue, Cambridge Biomedical Campus 3 , Cambridge, UK
4. Wellcome Centre for Cell Biology, Institute of Cell Biology, University of Edinburgh 4 , Edinburgh, UK