1. University of California, Irvine 1 Department of Developmental and Cell Biology, , Irvine, CA, USA
2. Ludwig Institute for Cancer Research 2 , La Jolla, CA, USA
3. University of California, San Diego 3 Department of Cell and Developmental Biology, , San Diego, CA, USA
4. University of California, San Diego 4 Department of Cellular and Molecular Medicine, , San Diego, CA, USA