1. School of Life Science and Technology, Tokyo Institute of Technology 1 , Yokohama, Japan
2. Faculty of Biology-Oriented Science and Technology, Kindai University 2 , Kinokawa, Japan
3. Graduate School of Agriculture, Kindai University 3 Department of Advanced Bioscience, , Nara, Japan
4. Cell Biology Center, Institute of Innovative Research, Tokyo Institute of Technology 4 , Yokohama, Japan
5. Institute for Quantitative Biosciences, The University of Tokyo 5 , Tokyo, Japan