1. Department of Pharmacology, Kyoto University Faculty of Medicine, Kyoto 606-8501, Japan
2. Horizontal Medical Research Organization, Kyoto University Faculty of Medicine, Kyoto 606-8501, Japan
3. Division of Cellular Therapy, Institute of Medical Science, University of Tokyo, Tokyo 108-8639, Japan
4. Core Research for Evolutionary Science and Technology Research Project, Kansai Advanced Research Center, Kobe 651-2492, Japan
5. Department of Biology, Graduate School of Science, Osaka University, Osaka 560-0043, Japan