1. These authors contributed equally to this work
2. Mineralized Tissue Section, The Hospital for Special Surgery, New York, New York, USA
3. Present Address: State University of New York at Stony Brook Medical School, Stony Brook, New York, USA
4. Present Address: New York University School of Medicine, New York, New York, USA
5. Present Address: New York Medical College, Valhalla, New York, USA
6. Weill Medical College of Cornell University, New York, New York, USA
7. Current address: University of Wisconsin Medical School, Madison, Wisconsin, USA