1. Department of Systems Biology, Yonsei University, Seoul 03722, Korea
2. These authors contributed equally to this work.
3. Platform Technology Research Center, Corporate R&D, LG Chem/LG Science Park, Seoul 07796, Korea
4. Gregor Mendel Institute (GMI), Austrian Academy of Sciences, Vienna BioCenter (VBC), 1030 Vienna, Austria
5. Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Incheon 22012, Korea
6. Department of Biology, Kyung Hee University, Seoul 02447, Korea