1. Department of Integrated Biological Sciences, Pusan National University, Busan 46241, Korea
2. Institute of Systems Biology, Pusan National University, Busan 46241, Korea
3. These authors contributed equally to this work.
4. Division of Life Sciences, Korea University, Seoul 02841, Korea
5. Department of Biological Sciences, College of Natural Sciences, Pusan National University, Busan 46231, Korea