1. Department of Biomedical Science, School of Medicine, Kyungpook National University, Daegu 41566, Korea
2. Department of Biochemistry and Cell Biology, School of Medicine, Kyungpook National University, Daegu 41566, Korea
3. These authors contributed equally to this work.
4. Department of Ophthalmology, Kyungpook National University Hospital, School of Medicine, Kyungpook National University, Daegu 41944, Korea