1. Department of Life Sciences, The Research Center for Cellular Homeostasis, Ewha Womans University, Seoul 03760, Korea
2. These authors contributed equally to this work.
3. Subtropical/Tropical Organism Gene Bank, Jeju National University, Jeju 63243, Korea
4. ExoMed, Inc., Seoul 01795, Korea
5. Department of Physiology, University of Ulsan College of Medicine, Asan Medical Center, Seoul 05505, Korea