1. Department of Medical Bioengineering, Graduate School of Natural Science and Technology, Okayama University, Okayama 700‑8530, Japan
2. Department of Oral Biochemistry, Asahi University School of Dentistry, Gifu 501‑0296, Japan
3. Department of Hematology and Oncology, Research Institute for Radiation Biology and Medicine, Hiroshima University, Hiroshima 734‑8553, Japan
4. Laboratory of Host Defenses, Mucosal Immunity Section, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, MD 20892, USA
5. Laboratory for Immune Regeneration, RIKEN Center for Integrative Medical Sciences, Yokohama 230‑0045, Japan
6. Department of Immunology, Institute for Frontier Medical Sciences, Kyoto University, Kyoto 606‑8507, Japan