1. Xiangya School of Medicine, Central South University, Changsha, Hunan 410083, P.R. China
2. Department of Cell Biology, School of Life Sciences, Central South University, Changsha, Hunan 410017, P.R. China
3. Hunan Provincial Key Laboratory of Nutrition and Quality Control of Aquatic Animals, Department of Biological and Environmental Engineering, Changsha University, Changsha, Hunan 410022, P.R. China