1. Laboratory of Zoology, Faculty of Science, University of Yaoundé I, PO Box 812, Yaoundé, Cameroon
2. Embrapa Florestas, Estrada da Ribeira, Km 111, Caixa postal 319, 83411-000, Colombo, PR, Brazil
3. Naturhistorisches Museum, Augustinergasse 2, CH-4001 Basel, Switzerland