1. Agriculture & Agri-Food Canada, Ottawa Research and Development Centre, 960 Carling Ave, Ottawa, ON K1A 0C6, Canada;
2. Muséum d'histoire naturelle de Genève, C.P. 6434, 1211 Genève 6, Suisse;
3. Systematic Entomology Laboratory, USDA, National Museum of National History, E-502, P.O. Box 37012, MRC 168, Washington, DC 20013-7012, United States of America;