1. School of Engineering, Newcastle University , Merz Court, Claremont Road , Newcastle Upon Tyne NE1 7RU , UK
2. LEPABE—Laboratory for Process Engineering, Environment, Biotechnology and Energy, Faculty of Engineering, University of Porto , Rua Dr. Roberto Frias, 4200-465 Porto , Portugal
3. Department of Life Sciences , Microbiology at Interfaces, Manchester Metropolitan University , Chester Street , Manchester M15GD , UK