1. Institute of Biology and Pathology , University Hospital of Grenoble and Alps , Grenoble , France
2. University Grenoble Alpes , Laboratory of Hypoxy Physiopathology Study Inserm U1042 , La Tronche , France
3. University Grenoble Alpes , DPM UMR 5063 , Grenoble , France
4. CNRS , DPM UMR 5063 , Grenoble , France