1. United Graduate School of Agricultural Science, Gifu University (Shizuoka University), 836 Ohya, Shizuoka 422Ð8529, Japan
2. Department of Applied Biological Chemistry, Faculty of Agriculture, Shizuoka University, Ohya, Shizuoka 422–8529, Japan
3. Basic Resarch Center, Shiseido Co. Ltd., Yokohama 224-8558, Japan