1. Biodiversity Research Group, Faculty of Science, Udon Thani Rajabhat University, Udon Thani 41000, Thailand
2. Walai Rukhavej Botanical Research Institute (WRBRI), Mahasarakham University, Maha Sarakham 44150, Thailand
3. Division of Environmental Health Sciences, Kochi Medical School, Kochi University, Oko, Nankoku 783-8505, Japan