1. Department of Biochemistry and Biotechnology , Laboratory of Plant and Environmental Biotechnology, University of Thessaly , Viopolis 41500 , Larissa , Greece
2. Department of Agricultural Development , Democritus University of Thrace , Orestiada , Greece
3. INRAE, Institut Agro, Univ. Bourgogne, Franche-Comté , Agroécologie Dijon , France