1. A.V. Zhirmunsky National Scientific Center of Marine Biology , Far Eastern Branch, Russian Academy of Sciences , Palchevskogo St., 17 , Vladivostok 690041 , Russia
2. Far Eastern State Technical Fisheries University , Lugovaya St., 52B , Vladivostok 690087 , Russia
3. Far Eastern Federal University , 10 Ajax Bay, Russky Island , Vladivostok 690922 , Russia