1. Department of Geoscience, University of Bremen , 28359 Bremen , Germany
2. Biogeochemistry and Geology Department, Leibniz Centre for Tropical Marine Research (ZMT) , 28359 Bremen , Germany
3. Oceanography and Hydrography Department, Kenya Marine and Fisheries Research Institute , 81651 – 80100 Mombasa , Kenya
4. Biology Department, Laboratory of Plant Biology and Nature Management (APNA), Vrije Universiteit Brussels , 1050 Brussels , Belgium