1. Department of Biological Sciences , 37580 National University of Singapore , Singapore 117558 , Singapore
2. Amakusa Marine Biological Laboratory , Kyushu University , Reihoku-Amakusa , Kumamoto 863-2507 , Japan
3. Lee Kong Chian Natural History Museum , 37580 National University of Singapore , Singapore 117377 , Singapore
4. Tropical Marine Science Institute , 37580 National University of Singapore , Singapore 119227 , Singapore
5. Centre for Nature-based Climate Solutions , 37580 National University of Singapore , Singapore 117558 , Singapore