1. Japan Agency for Marine-Earth Science and Technology , 2-15, Natsushima-cho , Yokosuka-city , Kanagawa 237-0061 , Japan
2. Environmental Dynamics and Management Group, Graduate School of Biosphere Science , Hiroshima University , 1-3-2 Kagamiyama , Higashi-Hiroshima City, Hiroshima 739-8511 , Japan
3. Division of Global Environmental Studies , Sophia University , 7-1 Kioicho, Chiyoda-ku , Tokyo 102-8554 , Japan
4. Department of Marine Science, Faculty of Fisheries , Kasetsart University , 50 Ngam Wong Wan Road, Ladyao Chatuchak , Bangkok 10900 , Thailand
5. Waterfront Vitalization and Environment Research Foundation , 405, Malissa Hills, 176-4, Asato, Naha , Okinawa 902-0067 , Japan
6. Akkeshi Marine Station, Field Science Center for Northern Biosphere , Hokkaido University , Aikappu 1, Akkeshi , Hokkaido 088-1113 , Japan