1. Institute of Evolutionary Ecology and Conservation Genomics, University of Ulm , Albert-Einstein-Allee 11, D-89069 Ulm , Germany
2. Smithsonian Tropical Research Institute , Apartado, 0843-03092 , Balboa Ancón , Panama
3. Ecology of Tropical Agricultural Systems , University of Hohenheim , Garbenstrasse 13, D-70599 Stuttgart , Germany