1. Teagasc, Environmental Research Centre, Johnstown Castle, Co. Wexford, Ireland
2. Agricultural Catchments Programme, Teagasc, Environmental Research Centre, Johnstown Castle, Co. Wexford, Ireland
3. Farming Systems Ecology, Wageningen University,Wageningen, The Netherlands
4. Soil Biology and Biological Soil Quality, Wageningen University, Wageningen, The Netherlands