1. Department of Oceanography , 37716 University of Cape Town , Cape Town , 7701 , South Africa
2. Elwandle Coastal Node , South African Environmental Observation Network (SAEON) , Gqeberha , 6001 , South Africa
3. Department of Biological Sciences , 37716 University of Cape Town , Rondebosch 7701 , South Africa
4. School of Biological and Chemical Sciences , University of Galway , University Road, H91TK33 Galway , Ireland
5. Department of Biodiversity and Conservation Biology , 56390 University of the Western Cape , Cape Town , 7535 , South Africa
6. Biodiversity & Conservation Biology Department , 56390 University of the Western Cape , Bellville , 7535 , South Africa