1. Research Center for Biosystematics and Evolution, National Research and Innovation Agency (BRIN), Cibinong 16911, Bogor, Indonesia
2. The Integrative Research Center, Field Museum of Natural History, Chicago, USA
3. Department of Biology, Faculty of Mathematics and Natural Sciences, Samudra University, Langsa, Aceh, Indonesia
4. Research Center for Ecology and Ethnobiology, National Research and Innovation Agency (BRIN), Cibinong 16911, Bogor, Indonesia