1. Department of Biology, School of Mathematics and Natural Sciences, Mongolian National University of Education, Ulaanbaatar, Mongolia
2. Climate Change Research and Cooperation Centre, Ministry of Environment and Tourism of Mongolia, Ulaanbaatar, Mongolia
3. Division of Biology, University of Montana, Missoula, Montanan, USA
4. Rhithron Associates, Inc., Missoula, Montana, USA
5. Institute for Mongolian Biodiversity and Ecological Studies Academy of Natural Sciences, Philadelphia, and Department of Biodiversity, Earth and Environmental Sciences, Drexel University, Philadelphia, Pennsylvania, USA