1. Study Program of Biotechnology, Faculty of Mathematics and Natural Sciences, Halu Oleo University, Kendari, Indonesia
2. Department of Biology, Faculty of Mathematics and Natural Sciences, Halu Oleo University, Kendari, Indonesia
3. Department of Publics Health, Faculty of Publics Health, Halu Oleo University, Kendari, Indonesia
4. Department of Natural Resources and Environmental Management, University of Hawaii at Manoa, Honolulu, HI, USA
5. Institute of Ocean and Earth Sciences, University of Malaya, Kuala Lumpur, Malaysia