1. a School of Biological Sciences , University of East Anglia , Norwich, NR4 7TJ, UK
2. b Animal Ecology Group, Centre for Ecological and Evolutionary Studies , University of Groningen , PO Box 14, 9750 AA Haren, The Netherlands
3. c Department of Marine Ecology , Royal Netherlands Institute for Sea Research (NIOZ) , PO Box 59, 1790 AB Den Burg, Texel, The Netherlands
4. d Conservation Science Group , University of Cambridge , Downing St., Cambridge, CB2 3EJ, UK