1. Department of Biology, Faculty of Mathematics and Natural Sciences, University of Sam Ratulangi, North Sulawesi, Indonesia
2. Department of Agricultural Technology, Faculty of Agriculture, University of Sam Ratulangi, North Sulawesi, Indonesia
3. Tasmanian Institute of Agriculture, The University of Tasmania, Hobart, TAS, Australia
4. School of Agriculture and Food Sustainability, The University of Queensland, Brisbane, QLD, Australia