1. Department of Otolaryngology, Head and Neck Surgery, Kyoto University Hospital/Graduate School of Medicine, Kyoto University, Kyoto, Japan;
2. Department of Physiology, Kurume University Hospital/Graduate School of Medicine, Kurume University, Kurume, Japan;
3. Institute of Biomedical Research and Innovation, Kobe, Japan;
4. Shiga Medical Center Research Institute, Shiga, Japan