1. Departments of Environmental Science and Sustainability, Biology, and Psychology, Drake University, Des Moines, IA, USA and Blank Park Zoo, Des Moines, IA, USA
2. Department of Environmental Science and Sustainability, Drake University, Des Moines, IA, USA
3. Veterinary Clinic, Blank Park Zoo, Des Moines, IA, USA and College of Veterinary Medicine, Iowa State University, Ames, IA, USA
4. Animal Care Department, Blank Park Zoo, Des Moines, IA, USA;