1. Seto Marine Biological Laboratory, Field Science Education and Research Center, Kyoto University, Wakayama, Japan
2. Wakayama, Japan
3. Department of Zoology, Division of Biological Science, Graduate School of Science, Kyoto University, Kyoto, Japan
4. CNRS, Univ. Littoral Côte d’Opale, UMR 8187, LOG, Laboratoire d’Océanologie et de Géosciences, Université de Lille, Lille, France
5. Department of Marine Resource and Energy, Tokyo University of Marine Science and Technology, Tokyo, Japan
6. Department of Zoology and Entomology, Rhodes University, Grahamstown, South Africa