1. Department of Biology, Faculty of Science, Gulu University, Gulu City, Uganda
2. The Regional Universities Forums for Capacity Building in Agriculture (RUFOUM), Makerere University, Kampala City, Uganda
3. College of Agriculture and Environmental Studies (CAES), Makerere University, Kampala City, Uganda
4. Centre for Geographic Information Systems (GIS) and Remote Sensing, Lund University, Lund City, Sweden
5. Departement of Soil Sciences, Université Evangélique en Afrique, Faculté d’Agronomie et Environnement, Bukavu, Democratic Republic of the Congo