1. Department of Genetics, Forestry and Agricultural, Biotechnology Institute (FABI), University of Pretoria, Pretoria, 0002, South Africa
2. Department of Microbiology and Plant Pathology, Forestry and Agricultural Biotechnology Institute, (FABI), University of Pretoria, Pretoria, 0002, South Africa
3. Department of Plant Pathology, Virginia Polytechnic Institute and State University, Blacksburg, Virginia 24061-0331