1. Nationaal Herbarium Nederland, Universiteit Leiden branch, Phanerogams and Cryptogams of the Netherlands and Europe section, P.O. Box 9514, 2300 RA Leiden, The Netherlands
2. Nationaal Herbarium Nederland, Universiteit Leiden branch, Molecular Systematics taskforce, P.O. Box 9514, 2300 RA Leiden, The Netherlands
3. Wageningen Agricultural University, Department of Environmental Sciences, Subdepartment of Soil Quality, P.O. Box 8005, 6700 EC Wageningen, The Netherlands