1. JST domestic fellow, Tohoku Research Center of Forestry and Forest Products Research Institute, Nabeyashiki 92-25, Shimo-Kuriyagawa, Morioka, Iwate 020-0123, Japan
2. Forestry and Agricultural Biotechnology Institute (FABI), Pretoria 0002, Republic of South Africa
3. Tohoku Research Center of Forestry and Forest Products Research Institute, Nabeyashiki 92-25, Shimo-Kuriyagawa, Morioka, Iwate 020-0123, Japan